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The Complete Oxidation of Glutamate Requires Metabolism of Part of the Carbon Skeleton via the Pyruvate Recycling Pathway Studies using 13 C-NMR spectroscopy have provided key insights into the metabolic fate of glutamate in astrocytes and information about the compartmentation of glutamate metabolism ( 13 C]glutamate into [U- 13 C]lactate (see Figure 1B) and also into [1,2- 13 C]glutamate and glutamine and specifically labeled molecules of aspartate confirms that the carbon skeleton of glutamate can be metabolized via the pyruvate recycling pathway in astrocytes and reenter the TCA cycle