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The average copper content in an adult human body is about 80mg, with the highest concentrations found in tissues such as the eyes, heart, liver, and brain ( Before extracellular Cu 2+ can be absorbed by cells, they need to be reduced to the monovalent reduced form of Cu + by metal reductases on the cell surface and then taken up by the cells through the solute carrier family 31 member 1 (SLC31A1, also known as CTR1) ( The copper within cells must be maintained at very low levels, with the vast majority bound to proteins ( 1+ in the cell are strong oxidants, catalyzing the generation of highly reactive carbonyl free radicals that can damage proteins, DNA, and lipids ( -15 M to 10 -21 M or less than one free copper ion per cell (58)